The Lions of the Triangle: An Update
Angama Mara
The clash with the Nyati males, the death of Slit Lip and the move east.
Coalition · Status uncertain
Also known as The Ngiro Are Males · Mara Triangle, Kenya
c. 2018 – present
Origin: Serengeti National Park, Tanzania (believed)
The Inselberg Males are a coalition of male lions in the Mara Triangle, the western part of Kenya’s Masai Mara. They take their name from the inselbergs, the isolated rocky hills near the border with Tanzania’s Serengeti where they first held territory, and are also known as the Ngiro Are or Ngiroare males. There were five of them: Manywele, Ruka, Nusu, Slit Lip and a fifth male usually called Olopolosie. Each is easily recognised by his scars: Slit Lip by a cleft upper lip, Nusu by a broken lower tooth, Ruka by torn ears.
Their origin is not certain. The Mara Predator Conservation Programme first recorded them in 2018 along the Kenya–Tanzania border on the Triangle side and believes they came from the neighbouring Serengeti; fans say they were first filmed around 2017 in the Lamai area of the northern Serengeti and were born around 2013–2014. By January 2020 all five were established in the south of the Triangle, where they took over prides including the Nasipa, Egyptian Goose and Purungat prides.
In late July 2022 Slit Lip, who often spent time alone with the prides, was caught and killed in a clash in which one of the young Nyati males also died. As the remaining four moved east towards Serena and Maji Machafu, the Nyati males took their old ground. By 2024 the Inselbergs were crossing the Mara River into the National Reserve, and in early 2025 Ruka was injured in a fight with the Kaskaz males from the Serengeti. The Purungat Pride’s young were all fathered by them. In 2026 fans reported that Manywele and Olopolosie had not been seen for many months; their fate has not been confirmed.
After the inselbergs, isolated rocky hills near the Kenya–Tanzania border in the south of the Mara Triangle, where they held territory.
Used by the Mara Predator Conservation Programme and some guides.
From ruruka, which Angama’s guides translate as “tear”, for the tears in his right ear.
Swahili for half, for the half-missing bottom right tooth.
For his split upper lip, possibly a birth defect.
The name later used for the fifth male, who was long unnamed. He is probably the male some reports call Ginger or Mkia; this is not confirmed.
Born, believed to be in the Serengeti; fans place them in the Lamai area of the north.
Reportedly filmed in Lamai, northern Serengeti, with the first pride they took over.
First recorded by the Mara Predator Conservation Programme on the Kenya side of the border in the Mara Triangle.
All five established in the south of the Triangle around the inselbergs.
Guides at Angama began naming them: Manywele, Ruka, Nusu, Slit Lip and an unnamed fifth male.
Ruka’s left eye was badly damaged, presumably in a fight with his brother Manywele.
Slit Lip was caught alone and killed by rival males, most likely the Nyati males, who lost one of their own in the clash.
The four survivors moved east, establishing themselves from Serena’s Sundowner Hill to Maji Machafu and coming into conflict with the Serena males.
All four were reported back in the Triangle, Manywele limping. They began crossing the Mara River into the National Reserve around Lookout.
Ruka was injured in a fight with the Kaskaz males from the Serengeti. The Purungat Pride’s sub-adults, all fathered by the Inselbergs, were noted to be almost maneless.
Fans reported that Manywele and Olopolosie had not been seen for many months, leaving Ruka and Nusu. Not confirmed.
Manywele of the Inselbergs is not the same lion as Manywele, the young male who rules the Black Rock Pride with Oloiborr. The two share a name, and both have lived in the Mara, which causes frequent confusion.
Angama’s guides describe a clash between the Inselbergs and the Nyati males in which one male from each side died. Some fans thought the Bila Shaka males might have been responsible. Both coalitions were in the area at the time.
The fifth Inselberg male was unnamed for years. Later sources call him Olopolosie; others list a Ginger or a Mkia in the coalition. They are probably the same lion, but this has not been confirmed.
The Inselbergs are now about twelve, old for male lions. Their status is uncertain, and this record will be updated.
Angama Mara
The clash with the Nyati males, the death of Slit Lip and the move east.
Mara Predator Conservation Programme
Field updates on the Inselberg males and the Purungat Pride.
YouTube
The four surviving brothers together after the death of Slit Lip.
Where sources disagree, the most widely reported figure is shown, with other reports noted beneath it.